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There are several factors that could have affected this result. This could include the chemical composition of the soil, water availability and even the weather! There could also be increased competition for resources in one area more than another, both inter-species and intra-species. You could also think about the likelihood of one area being more polluted than another.

The upturned soil area would have provided more shelter for the invertebrates than the open grassland, protecting then from extreme wether conditions and provided more areas to hide from predators. It also would have retained water more effectively than the open grassland, which is very important for the survival of invertebrates that dry out easily such as earthworms and snails. This means that the species colonising this area do not need to be as specialised to the environment as in the grassland, providing a wider range of niches to be occupied in less (evolutionary) time. Hope this helps!

apart from light intensity you should also think about predators more in a certain area or less protective places to hide. in a certain area

Furthermore food and water availability. Shelter from the elements. suitability of nesting site/places to lay eggs etc. 

hope that helps

In addition to light intensity there are a number of factors that could affect this:

Availability of food/water, shelter from the elements, shelter from predators, presence of any symbiotic organisms and presence of possible mates.

Hope this helps a bit :)

This link might help you, even though it applies to beaches it's still relevant :)

Availability of food (soil richness) shelter, water availability. What was the hypothesis etc? Might help me help you a little more to know some more background info :)

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